PenumbraPEN
13F holders, Q1 2024 (March 31)342holders
6famous investors
$223.13avg. 13F price*
At the end of Q1 2024, 342 institutions reported holding Penumbra Inc. (PEN) in their 13F filings. The largest holders were FMR LLC ($1.3B), BlackRock Inc. ($906.4M) and Vanguard Group Inc. ($823.7M). 6 famous investors held it, including Ray Dalio (0.0% of Bridgewater Associates's portfolio), Israel Englander (0.0% of Millennium Management's portfolio) and Joel Greenblatt (0.0% of Gotham Asset Management's portfolio).
- Company
- Penumbra Inc.
- CUSIP
- 70975L107
- Latest filing quarter
- Q2 2026
History
Number of 13F holders
Total Exit/New Position Q1 2024
| Institutions | Total Value | |
|---|---|---|
| New Positions | 45 | $98.7M |
| Exits | 42 | $44.6M |
Ownership 342 holders
| FMR LLC | 0.09% | Add +0.3% | 5,805,506 | $1.3B |
| BlackRock Inc. | 0.02% | Add +1.5% | 4,061,245 | $906.4M |
| Vanguard Group Inc. | 0.02% | Add +0.9% | 3,690,660 | $823.7M |
| T. Rowe Price Associates Inc. | 0.08% | Reduce -2.5% | 3,025,823 | $675.3M |
| State Street Corp. | 0.01% | Add +9.4% | 1,204,327 | $268.8M |
| Capital Research Global Investors | 0.06% | Reduce -57.8% | 1,130,866 | $252.4M |
| Champlain Investment Partners, LLC | 1.39% | Add +131.8% | 1,084,595 | $242.1M |
| William Blair Investment Management, LLC | 0.48% | Add +12.2% | 833,703 | $186.1M |
| Clearbridge Investments, LLC | 0.14% | Reduce -1.8% | 786,336 | $175.5M |
| RTW Investments, LP | 2.23% | Add +2.0% | 675,888 | $150.8M |
| Baillie Gifford & Co. | 0.11% | Reduce -2.1% | 634,319 | $141.6M |
| Geode Capital Management, LLC | 0.01% | Add +4.0% | 615,849 | $137.5M |
| Capital World Investors | 0.02% | No change | 543,274 | $121.2M |
| Rock Springs Capital Management LP | 2.87% | Reduce -6.0% | 505,000 | $112.7M |
| Morgan Stanley | 0.01% | Add +6.6% | 389,056 | $86.8M |
| Eagle Asset Management Inc. | 0.38% | Reduce -1.3% | 345,551 | $75.1M |
| Bank of New York Mellon Corp | 0.01% | Reduce -4.3% | 322,498 | $72.0M |
| Braidwell LP | 3.72% | Add +32.8% | 295,892 | $66.0M |
| Norges Bank | 0.01% | Reduce -13.7% | 292,368 | $65.3M |
| Franklin Resources Inc. | 0.02% | Add +17.4% | 265,666 | $59.3M |
| Charles Schwab Investment Management Inc. | 0.01% | Add +2.2% | 255,520 | $57.0M |
| Janus Henderson Group PLC | 0.03% | Reduce -1.0% | 254,802 | $56.9M |
| Northern Trust Corp. | 0.01% | Reduce -12.6% | 237,587 | $53.0M |
| Allspring Global Investments Holdings, LLC | 0.07% | Reduce -28.9% | 188,399 | $42.0M |
| Bellevue Group AG | 0.62% | Reduce -10.8% | 181,158 | $40.4M |
| Baird Financial Group, Inc. | 0.08% | New | 171,861 | $38.4M |
| Alliancebernstein L.P. | 0.01% | Add +16.4% | 171,124 | $38.2M |
| American Century Companies Inc. | 0.02% | Reduce -0.6% | 168,097 | $37.5M |
| Nuveen Asset Management, LLC | 0.01% | Reduce -8.3% | 161,574 | $36.1M |
| PFM Health Sciences, LP | 1.77% | No change | 155,786 | $34.8M |
| Principal Financial Group Inc. | 0.02% | Add +1.9% | 145,741 | $32.5M |
| Artisan Partners Limited Partnership | 0.05% | Reduce -4.9% | 139,087 | $31.0M |
| Dimensional Fund Advisors LP | 0.01% | Add +10.0% | 130,666 | $29.2M |
| TD Asset Management Inc | 0.03% | Reduce -4.9% | 129,786 | $29.0M |
| Qube Research & Technologies Ltd | 0.05% | Add +60.1% | 129,095 | $28.8M |
| Balyasny Asset Management L.P. | 0.07% | Add +329.9% | 126,261 | $28.2M |
| Polar Capital Holdings Plc | 0.13% | Reduce -11.8% | 112,452 | $25.1M |
| Rhumbline Advisers | 0.02% | Reduce -3.6% | 108,615 | $24.2M |
| AXA S.A. | 0.07% | Add +8.3% | 105,535 | $23.6M |
| Manufacturers Life Insurance Company, The | 0.02% | Add +3.3% | 103,132 | $23.0M |
| JPMorgan Chase & Co. | 0.00% | Add +40.6% | 102,610 | $22.9M |
| Loomis Sayles & Co. L P | 0.03% | Add +10.6% | 95,985 | $21.4M |
| T. Rowe Price Investment Management, Inc. | 0.01% | Add +41.9% | 95,522 | $21.3M |
| Millennium Management | 0.02% | Reduce -22.4% | 94,116 | $21.0M |
| Man Group plc | 0.06% | Add +124.1% | 91,194 | $20.4M |
| Jennison Associates LLC | 0.01% | Add +192.1% | 87,139 | $19.4M |
| Goldman Sachs Group Inc. | 0.00% | Reduce -29.7% | 86,385 | $19.3M |
| Soleus Capital Management, L.P. | 1.68% | Add +7.9% | 85,700 | $19.1M |
| Amundi | 0.01% | Reduce -17.7% | 92,505 | $18.7M |
| Bank of America Corp. | 0.00% | Add +78.5% | 79,734 | $17.8M |
| Carret Asset Management, LLC | 2.08% | No change | 79,000 | $17.6M |
| C WorldWide Group Holding A/S | 0.20% | New | 77,000 | $17.2M |
| Swiss National Bank | 0.01% | Add +0.6% | 72,400 | $16.2M |
| Integral Health Asset Management, LLC | 1.53% | Add +75.0% | 70,000 | $15.6M |
| Twinbeech Capital LP | 0.31% | New | 67,759 | $15.1M |
| UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC | 0.01% | Add +9.4% | 67,491 | $15.1M |
| Invesco Ltd. | 0.00% | Reduce -83.3% | 63,051 | $14.1M |
| Azimuth Capital Investment Management LLC | 0.58% | Reduce -5.4% | 62,650 | $14.0M |
| Legal & General Group Plc | 0.00% | Add +0.3% | 61,803 | $13.8M |
| Federated Hermes, Inc. | 0.03% | Reduce -14.8% | 61,750 | $13.8M |
| Wells Fargo & Company | 0.00% | Add +5.2% | 58,980 | $13.2M |
| Ameriprise Financial Inc. | 0.00% | Reduce -1.4% | 55,011 | $12.3M |
| Credit Suisse AG/ | 0.01% | Add +1.3% | 53,623 | $12.0M |
| New York State Common Retirement Fund | 0.02% | Reduce -4.2% | 52,646 | $11.7M |
| State Board of Administration of Florida Retirement System | 0.03% | Add +7.2% | 41,548 | $11.6M |
| American International Group, Inc. | 0.04% | Reduce -2.0% | 51,377 | $11.5M |
| California Public Employees Retirement System | 0.01% | Reduce -23.0% | 50,078 | $11.2M |
| Squarepoint Ops LLC | 0.05% | Add +2212.8% | 49,448 | $11.0M |
| Retirement Systems of Alabama | 0.04% | Add +0.3% | 48,467 | $10.8M |
| Rhenman & Partners Asset Management AB | 0.97% | Add +1740.0% | 46,000 | $10.3M |
| State of Wisconsin Investment Board | 0.03% | Reduce -9.5% | 45,707 | $10.2M |
| Redmile Group, LLC | 0.37% | Reduce -53.2% | 44,613 | $10.0M |
| Primecap Management Co. | 0.01% | No change | 42,860 | $9.6M |
| Citadel Advisors | 0.01% | Reduce -53.3% | 42,715 | $9.5M |
| Jane Street Group, LLC | 0.01% | Reduce -50.2% | 41,573 | $9.3M |
| Sei Investments Co. | 0.01% | Add +62.5% | 39,505 | $8.8M |
| New York State Teachers Retirement System | 0.02% | Add +1.1% | 38,288 | $8.5M |
| First Light Asset Management, LLC | 0.65% | New | 37,955 | $8.5M |
| Barclays PLC | 0.00% | Add +12.0% | 37,766 | $8.4M |
| California State Teachers Retirement System | 0.01% | Reduce -5.0% | 36,668 | $8.2M |
| Cordgrass Capital Advisors, LLC | 2.40% | Add +0.7% | 34,607 | $7.7M |
| Texas Permanent School Fund Corp. | 0.06% | Add +1.3% | 33,779 | $7.5M |
| AQR Capital Management | 0.01% | Add +3234.9% | 33,049 | $7.4M |
| Atika Capital Management LLC | 0.83% | No change | 33,000 | $7.4M |
| CANADA LIFE ASSURANCE Co | 0.01% | Add +2.0% | 29,054 | $6.5M |
| Teachers Retirement System of the State of Kentucky | 0.06% | Reduce -1.2% | 28,746 | $6.4M |
| Laird Norton Wetherby Wealth Management, LLC | 0.29% | Add +0.0% | 26,839 | $6.0M |
| Congress Asset Management Co. | 0.04% | Add +375.0% | 26,576 | $5.9M |
| Aquatic Capital Management LLC | 0.19% | Add +215.5% | 26,500 | $5.9M |
| Group One Trading LLC | 0.20% | Add +66.2% | 25,152 | $5.6M |
| Mutual of America Capital Management LLC | 0.06% | Reduce -3.3% | 23,921 | $5.3M |
| Bank of Montreal | 0.00% | Add +31.5% | 23,246 | $5.3M |
| Trillium Asset Management, LLC | 0.13% | Add +2.0% | 23,401 | $5.2M |
| MetLife Investment Management, LLC | 0.03% | Reduce -2.4% | 22,658 | $5.1M |
| Voya Investment Management LLC | 0.01% | Add +8.2% | 22,102 | $4.9M |
| Point72 Hong Kong Ltd | 0.24% | New | 21,425 | $4.8M |
| Amalgamated Bank | 0.04% | Reduce -18.3% | 21,318 | $4.8M |
| Next Century Growth Investors LLC | 0.41% | Reduce -10.4% | 21,290 | $4.8M |
| State of New Jersey Common Pension Fund D | 0.01% | No change | 20,733 | $4.6M |
| Marshall Wace, LLP | 0.01% | Reduce -59.7% | 20,435 | $4.6M |
| Bridgewater Associates | 0.02% | Reduce -47.7% | 20,364 | $4.5M |
| Public Employees Retirement System of Ohio | 0.02% | Reduce -16.5% | 20,232 | $4.5M |
| ExodusPoint Capital Management, LP | 0.06% | Reduce -31.6% | 20,060 | $4.5M |
| Envestnet Asset Management Inc. | 0.00% | Add +80.6% | 18,864 | $4.2M |
| EULAV Asset Management | 0.11% | No change | 18,100 | $4.0M |
| Yousif Capital Management, LLC | 0.04% | Reduce -2.5% | 17,941 | $4.0M |
| Blair William & Co. | 0.01% | Add +24.3% | 16,674 | $3.7M |
| Beacon Pointe Advisors, LLC | 0.04% | Reduce -30.1% | 16,443 | $3.7M |
| Treasurer of the State of North Carolina | 0.02% | Reduce -2.2% | 16,199 | $3.6M |
| Candriam S.C.A. | 0.02% | Reduce -47.1% | 16,171 | $3.6M |
| Engineers Gate Manager LP | 0.09% | Add +1156.6% | 15,343 | $3.4M |
| E. Ohman J:or Asset Management AB | 0.09% | Reduce -31.9% | 15,312 | $3.4M |
| Comerica Bank | 0.01% | Reduce -3.3% | 15,308 | $3.4M |
| State of Tennessee, Department of Treasury | 0.01% | Reduce -6.0% | 15,000 | $3.3M |
| Polar Asset Management Partners Inc. | 0.12% | Add +23.3% | 14,761 | $3.3M |
| UBS Group AG | 0.00% | Add +58.8% | 14,630 | $3.3M |
| Raymond James & Associates | 0.00% | Add +32.6% | 14,076 | $3.1M |
| Trexquant Investment LP | 0.06% | Add +28.8% | 12,963 | $2.9M |
| PEAK6 Investments LLC | 0.14% | Add +76.2% | 12,882 | $2.9M |
| Ensign Peak Advisors, Inc | 0.01% | Add +4.6% | 12,671 | $2.8M |
| Neuberger Berman Group LLC | 0.00% | Reduce -80.3% | 12,269 | $2.7M |
| LMR Partners LLP | 0.10% | Add +41.4% | 11,992 | $2.7M |
| Deutsche Bank AG | 0.00% | Reduce -62.3% | 11,973 | $2.7M |
| Victory Capital Management Inc. | 0.00% | Reduce -63.7% | 11,368 | $2.5M |
| CW Advisors, LLC | 0.04% | Add +111.5% | 11,356 | $2.5M |
| LPL Financial LLC | 0.00% | Add +76.3% | 11,231 | $2.5M |
| Louisiana State Employees Retirement System | 0.05% | Add +2.8% | 11,100 | $2.5M |
| Illinois Municipal Retirement Fund | 0.04% | No change | 10,793 | $2.4M |
| US Bancorp \de | 0.00% | Add +5.3% | 10,689 | $2.4M |
| Canada Pension Plan Investment Board | 0.00% | Reduce -75.4% | 10,600 | $2.4M |
| Thoroughbred Financial Services, LLC | 0.22% | Reduce -11.0% | 10,239 | $2.3M |
| Algert Global LLC | 0.09% | Reduce -3.1% | 10,150 | $2.3M |
| Arizona State Retirement System | 0.02% | Reduce -1.3% | 10,077 | $2.2M |
| Centiva Capital, LP | 0.14% | Add +22.1% | 10,075 | $2.2M |
| Verition Fund Management LLC | 0.02% | Add +626.9% | 9,813 | $2.2M |
| Mitsubishi UFJ Asset Management Co., Ltd. | 0.00% | Reduce -8.0% | 9,698 | $2.2M |
| Thrivent Financial for Lutherans | 0.00% | Add +0.3% | 9,672 | $2.2M |
| Oregon Public Employees Retirement Fund | 0.04% | Reduce -12.4% | 9,541 | $2.1M |
| State of Michigan Retirement System | 0.01% | Add +3.3% | 9,514 | $2.1M |
| Guggenheim Capital LLC | 0.01% | Add +332.3% | 9,304 | $2.1M |
| Jump Financial, LLC | 0.06% | Add +160.5% | 9,118 | $2.0M |
| Handelsbanken Fonder AB | 0.01% | No change | 9,058 | $2.0M |
| Mariner, LLC | 0.00% | Add +0.6% | 8,689 | $1.9M |
| Northwestern Mutual Investment Management Company, LLC | 0.05% | Add +0.3% | 8,609 | $1.9M |
| Commonwealth of Pennsylvania Public School Empls Retrmt SYS | 0.01% | Add +0.3% | 8,538 | $1.9M |
| Ameritas Investment Partners, Inc. | 0.07% | No change | 8,522 | $1.9M |
| Xponance LLC | 0.02% | Add +1.3% | 8,259 | $1.8M |
| AlTi Global, Inc. | 0.05% | No change | 7,804 | $1.7M |
| New Mexico Educational Retirement Board | 0.06% | No change | 7,800 | $1.7M |
| ClariVest Asset Management LLC | 0.17% | Reduce -0.9% | 7,504 | $1.7M |
| Exome Asset Management LLC | 1.41% | Reduce -14.0% | 7,353 | $1.6M |
| Genus Capital Management Inc. | 0.21% | — | 7,300 | $1.6M |
| Teacher Retirement System of Texas | 0.01% | Reduce -18.5% | 7,149 | $1.6M |
| XTX Topco Ltd | 0.21% | Add +768.9% | 6,977 | $1.6M |
| Motley Fool Asset Management LLC | 0.10% | Reduce -0.9% | 6,967 | $1.6M |
| Y-Intercept (Hong Kong) Ltd | 0.11% | New | 6,825 | $1.5M |
| Sumitomo Mitsui Trust Group, Inc. | 0.00% | Add +0.4% | 6,793 | $1.5M |
| Diversified Trust Co. | 0.05% | Add +112.1% | 6,720 | $1.5M |
| Dynamic Technology Lab Private Ltd | 0.16% | Add +41.0% | 6,703 | $1.5M |
| BNP Paribas Financial Markets | 0.00% | Reduce -66.0% | 6,673 | $1.5M |
| Utah Retirement Systems | 0.02% | Reduce -0.7% | 6,317 | $1.4M |
| Allianz Asset Management GmbH | 0.00% | Reduce -12.5% | 6,282 | $1.4M |
| Zurcher Kantonalbank (Zurich Cantonalbank) | 0.00% | Reduce -5.6% | 6,272 | $1.4M |
| Oak Ridge Investments LLC | 0.14% | Reduce -4.8% | 6,271 | $1.4M |
| Bessemer Group Inc. | 0.00% | Reduce -30.6% | 6,229 | $1.4M |
| Fox Run Management, L.L.C. | 0.30% | Add +113.2% | 6,162 | $1.4M |
| Pacer Advisors, Inc. | 0.00% | Reduce -27.2% | 5,872 | $1.3M |
| OCONNOR, A Distinct Business Unit of UBS ASSET MANAGEMENT AMERICAS (LLC) | 0.04% | Add +2.7% | 5,798 | $1.3M |
| Linden Thomas Advisory Services, LLC | 0.21% | Add +2.0% | 5,788 | $1.3M |
| Cubist Systematic Strategies, LLC | 0.01% | New | 5,714 | $1.3M |
| Russell Investments Group, Ltd. | 0.00% | Reduce -57.5% | 5,400 | $1.2M |
| Walleye Capital LLC | 0.01% | Add +121.2% | 5,344 | $1.2M |
| Summit Trail Advisors, LLC | 0.03% | Reduce -0.2% | 5,343 | $1.2M |
| Tower Research Capital LLC (TRC) | 0.03% | Add +27.8% | 5,339 | $1.2M |
| Townsquare Capital LLC | 0.02% | New | 5,138 | $1.1M |
| Washington Trust Advisors, Inc. | 0.10% | No change | 4,996 | $1.1M |
| Tidal Investments LLC | 0.02% | Add +276.9% | 4,971 | $1.1M |
| Gotham Asset Management | 0.02% | Add +509.3% | 4,868 | $1.1M |
| Highland Capital Management, LLC | 0.06% | Add +0.2% | 4,777 | $1.1M |
| Exchange Traded Concepts, LLC | 0.03% | Add +0.2% | 4,624 | $1.0M |
| Pictet Asset Management Holding SA | 0.00% | Add +8.3% | 4,593 | $1.0M |
| New York Life Investment Management LLC | 0.01% | Reduce -1.9% | 4,274 | $954K |
| State of Alaska, Department of Revenue | 0.01% | Add +0.4% | 4,185 | $934K |
| Graham Capital Management, L.P. | 0.05% | Reduce -39.5% | 4,141 | $924K |
| Vanguard Personalized Indexing Management, LLC | 0.02% | Reduce -23.6% | 4,135 | $923K |
| Public Employees Retirement Association of Colorado | 0.00% | Reduce -6.8% | 4,071 | $909K |
| Virtu Financial LLC | 0.07% | Add +61.6% | 4,011 | $895K |
| S&co Inc. | 0.06% | New | 4,000 | $892K |
| Employees Retirement System of Texas | 0.01% | Add +107.5% | 3,999 | $892K |
| Lecap Asset Management Ltd. | 0.18% | Add +217.9% | 3,951 | $882K |
| Rwa Wealth Partners, LLC | 0.01% | No change | 3,825 | $854K |
| Two Sigma Investments, LP | 0.00% | Add +76.2% | 3,701 | $826K |
| Prelude Capital Management, LLC | 0.06% | Add +146.1% | 3,686 | $823K |
| Citigroup Inc. | 0.00% | Reduce -15.9% | 3,598 | $803K |
| Pathstone Holdings, LLC | 0.00% | Add +13.0% | 3,467 | $774K |
| Wolverine Trading, LLC | 0.01% | Add +29.1% | 3,370 | $752K |
| Point72 (DIFC) Ltd | 0.06% | — | 3,313 | $739K |
| Onyx Bridge Wealth Group LLC | 0.13% | No change | 3,575 | $731K |
| Baker Avenue Asset Management, LP | 0.03% | Reduce -0.3% | 3,272 | $730K |
| Inceptionr LLC | 0.19% | Add +256.2% | 3,231 | $721K |
| Securian Asset Management, Inc. | 0.02% | Reduce -3.1% | 3,155 | $704K |
| Van Eck Associates Corp. | 0.00% | New | 3,129 | $698K |
| Cinctive Capital Management LP | 0.04% | New | 3,118 | $696K |
| Janney Montgomery Scott LLC | 0.00% | New | 3,088 | $689K |
| Seven Eight Capital, LP | 0.07% | New | 3,046 | $680K |
| Rathbones Group PLC | 0.00% | No change | 3,000 | $670K |
| Scientech Research LLC | 0.34% | New | 2,914 | $650K |
| Leavell Investment Management, Inc. | 0.04% | Reduce -27.5% | 2,900 | $647K |
| Crossmark Global Holdings, Inc. | 0.01% | Add +2.1% | 2,872 | $641K |
| Corient Private Wealth LP | 0.00% | Reduce -23.0% | 2,861 | $639K |
| Vestcor Inc | 0.02% | New | 2,806 | $626K |
| HSBC Holdings PLC | 0.00% | Reduce -14.9% | 2,766 | $617K |
| Maryland State Retirement & Pension System | 0.01% | No change | 2,765 | $580K |
| Prudential Financial Inc. | 0.00% | Reduce -6.7% | 2,567 | $573K |
| QRG Capital Management, Inc. | 0.01% | Reduce -18.4% | 2,547 | $568K |
| VCI Wealth Management LLC | 0.38% | New | 2,487 | $555K |
| PNC Financial Services Group, Inc. | 0.00% | Add +45.2% | 2,371 | $529K |
| Royal Bank of Canada | 0.00% | Reduce -6.0% | 2,298 | $514K |
| Abbot Financial Management, Inc. | 0.28% | Add +7.5% | 2,238 | $499K |
| Seaport Global Advisors, LLC | 1.01% | Reduce -34.4% | 2,225 | $497K |
| Susquehanna Portfolio Strategies, LLC | 0.01% | Reduce -88.9% | 2,167 | $484K |
| Blueshift Asset Management, LLC | 0.25% | New | 2,157 | $481K |
| HighTower Advisors, LLC | 0.00% | Add +7.8% | 2,122 | $473K |
| ProShare Advisors LLC | 0.00% | Reduce -8.5% | 2,109 | $471K |
| Olympiad Research LP | 0.30% | Add +69.9% | 2,069 | $462K |
| Mercer Global Advisors Inc. | 0.00% | Add +16.0% | 2,063 | $460K |
| Two Sigma Securities, LLC | 0.02% | New | 1,931 | $431K |
| Yorktown Management & Research Co Inc | 0.39% | No change | 1,850 | $413K |
| Pearl River Capital, LLC | 0.06% | New | 1,837 | $410K |
| Axq Capital, LP | 0.10% | New | 1,820 | $406K |
| Gsa Capital Partners LLP | 0.03% | Add +35.6% | 1,792 | $400K |
| Quent Capital, LLC | 0.05% | Add +17660.0% | 1,776 | $396K |
| Walleye Trading LLC | 0.01% | Add +33.9% | 1,753 | $391K |
| Campbell & CO Investment Adviser LLC | 0.03% | New | 1,722 | $384K |
| Silvercrest Asset Management Group LLC | 0.00% | Add +3.4% | 1,680 | $375K |
| Shelton Capital Management | 0.01% | Add +86.9% | 1,630 | $364K |
| D. E. Shaw & Co. | 0.00% | New | 1,559 | $348K |
| Profund Advisors LLC | 0.01% | Add +21.9% | 1,528 | $341K |
| PDT Partners, LLC | 0.03% | Reduce -1.0% | 1,521 | $339K |
| Norden Group LLC | 0.07% | New | 1,438 | $321K |
| NEOS Investment Management LLC | 0.02% | New | 1,433 | $320K |
| Cynosure Group, LLC | 0.06% | New | 1,427 | $318K |
| M&T Bank Corp | 0.00% | Add +2.5% | 1,414 | $316K |
| Alliance Wealth Advisors, LLC | 0.09% | New | 1,331 | $297K |
| Bridgefront Capital, LLC | 0.22% | New | 1,285 | $287K |
| 17 Capital Partners, LLC | 0.21% | No change | 1,281 | $286K |
| HRT Financial LP | 0.00% | Reduce -74.0% | 1,244 | $277K |
| Aigen Investment Management, LP | 0.06% | Reduce -49.4% | 1,238 | $276K |
| Prospera Financial Services Inc | 0.01% | Reduce -12.6% | 1,180 | $263K |
| Concorde Asset Management, LLC | 0.18% | No change | 1,175 | $262K |
| Creative Planning | 0.00% | Add +14.8% | 1,155 | $258K |
| Cutler Group LLC / CA | 0.12% | New | 1,100 | $245K |
| Envestnet Portfolio Solutions, Inc. | 0.00% | New | 1,082 | $241K |
| Coldstream Capital Management Inc. | 0.01% | Add +4.0% | 1,037 | $231K |
| CI Investments Inc. | 0.00% | Add +80.4% | 1,023 | $228K |
| Family Capital Management, Inc. | 0.16% | No change | 1,000 | $223K |
| CIBC Asset Management Inc | 0.00% | No change | 994 | $222K |
| Petrus Trust Company, LTA | 0.02% | New | 990 | $221K |
| Meiji Yasuda Asset Management Co Ltd. | 0.01% | New | 989 | $221K |
| Simplex Trading, LLC | 0.01% | Reduce -88.1% | 989 | $220K |
| Integrated Wealth Concepts LLC | 0.00% | Add +2.3% | 947 | $211K |
| Vident Advisory, LLC | 0.00% | Reduce -2.3% | 945 | $211K |
| Arcadia Investment Management Corp. | 0.04% | Add +902.1% | 932 | $208K |
| Artemis Investment Management LLP | 0.00% | Reduce -59.0% | 930 | $207K |
| Cetera Investment Advisers | 0.00% | New | 911 | $203K |
| Shell Asset Management Co. | 0.01% | Reduce -9.1% | 1,068 | $192K |
| AlphaQuest LLC | 0.03% | Add +186.9% | 812 | $181K |
| Leonteq Securities AG | 0.02% | Add +24.6% | 791 | $177K |
| KBC Group NV | 0.00% | Add +12.3% | 783 | $175K |
| Quadrant Capital Group LLC | 0.01% | Reduce -3.7% | 695 | $155K |
| Lazard Asset Management LLC | 0.00% | Reduce -2.1% | 548 | $122K |
| Covestor Ltd | 0.08% | Add +1.1% | 537 | $120K |
| EntryPoint Capital, LLC | 0.10% | Add +562.8% | 517 | $115K |
| RiverPark Advisors, LLC | 0.09% | Add +360.9% | 507 | $113K |
| Cwm, LLC | 0.00% | Reduce -3.9% | 491 | $110K |
| Osaic Holdings, Inc. | 0.00% | Reduce -81.7% | 462 | $103K |
| Nisa Investment Advisors, LLC | 0.00% | Add +15.4% | 413 | $92K |
| Fifth Third Bancorp | 0.00% | Add +7.4% | 406 | $91K |
| Headlands Technologies LLC | 0.02% | New | 398 | $89K |
| Federation des caisses Desjardins du Quebec | 0.00% | Reduce -99.3% | 378 | $84K |
| Advisors Asset Management, Inc. | 0.00% | Reduce -18.3% | 340 | $76K |
| Signaturefd, LLC | 0.00% | Add +11.2% | 239 | $53K |
| Quarry LP | 0.02% | Add +340.0% | 220 | $49K |
| Allworth Financial LP | 0.00% | Add +397.0% | 164 | $37K |
| CIBC Private Wealth Group LLC | 0.00% | New | 167 | $36K |
| Mather Group, Llc. | 0.00% | New | 132 | $33K |
| Steward Partners Investment Advisory, LLC | 0.00% | Reduce -3.5% | 140 | $31K |
| Parallel Advisors, LLC | 0.00% | Add +9.7% | 125 | $28K |
| Huntington National Bank | 0.00% | Reduce -65.7% | 123 | $27K |
| Principal Securities, Inc. | 0.00% | Add +74.6% | 110 | $25K |
| Benjamin F. Edwards & Company, Inc. | 0.00% | Add +1.9% | 108 | $24K |
| MassMutual Private Wealth & Trust, FSB | 0.00% | Add +4.2% | 99 | $22K |
| Global Retirement Partners, LLC | 0.00% | Add +11.9% | 94 | $21K |
| GAMMA Investing LLC | 0.00% | Add +9.9% | 89 | $20K |
| Annis Gardner Whiting Capital Advisors, LLC | 0.01% | New | 73 | $16K |
| Beacon Capital Management, LLC | 0.00% | Reduce -26.3% | 59 | $13K |
| Lindbrook Capital, LLC | 0.00% | Reduce -9.1% | 50 | $11K |
| Sandy Spring Bank | 0.00% | New | 50 | $11K |
| Lanark Financial, Inc. | 0.00% | No change | 45 | $11K |
| Farther Finance Advisors, LLC | 0.00% | Add +2.7% | 38 | $8K |
| First Manhattan Co. Llc. | 0.00% | New | 37 | $8K |
| LGT Financial Advisors LLC | 0.01% | No change | 27 | $6K |
| MCF Advisors LLC | 0.00% | No change | 24 | $5K |
| First Horizon Advisors, Inc. | 0.00% | No change | 22 | $5K |
| Sunbelt Securities, Inc. | 0.00% | No change | 21 | $5K |
| Key Financial Inc | 0.00% | No change | 20 | $4K |
| Northwestern Mutual Wealth Management Co. | 0.00% | Add +100.0% | 20 | $4K |
| Meeder Asset Management Inc. | 0.00% | New | 18 | $4K |
| Harbour Investments, Inc. | 0.00% | Add +125.0% | 18 | $4K |
| Valley National Advisers Inc. | 0.00% | No change | 21 | $4K |
| Smithfield Trust Co | 0.00% | No change | 17 | $4K |
| CoreCap Advisors, LLC | 0.00% | New | 17 | $4K |
| Farmers & Merchants Investments Inc. | 0.00% | No change | 16 | $4K |
| CVA Family Office, LLC | 0.00% | No change | 15 | $3K |
| EverSource Wealth Advisors, LLC | 0.00% | Add +66.7% | 15 | $3K |
| FinTrust Capital Advisors, LLC | 0.00% | New | 13 | $3K |
| UMB Bank, n.a. | 0.00% | New | 10 | $2K |
| Spire Wealth Management | 0.00% | No change | 10 | $2K |
| NewEdge Advisors, LLC | 0.00% | No change | 10 | $2K |
| Bell Investment Advisors, Inc | 0.00% | Add +28.6% | 9 | $2K |
| Avion Wealth | 0.00% | No change | 6 | $2K |
| Raffa Wealth Management, LLC | 0.00% | No change | 8 | $2K |
| IFP Advisors, Inc | 0.00% | Add +700.0% | 8 | $2K |
| Smartleaf Asset Management LLC | 0.00% | Add +600.0% | 7 | $2K |
| Bokf, NA | 0.00% | No change | 7 | $2K |
| SYSTM Wealth Solutions LLC | 0.00% | No change | 6 | $1K |
| Janiczek Wealth Management, LLC | 0.00% | No change | 5 | $1K |
| JFS Wealth Advisors, LLC | 0.00% | No change | 5 | $1K |
| Roble, Belko & Company, Inc | 0.00% | No change | 5 | $1K |
| Financial Management Professionals, Inc. | 0.00% | No change | 3 | $1K |
| Capital Advisors, Ltd. LLC | 0.00% | No change | 4 | $1K |
| Richardson Financial Services Inc. | 0.00% | New | 4 | $893 |
| Private Wealth Management Group, LLC | 0.00% | No change | 4 | $893 |
| Riggs Asset Managment Co. Inc. | 0.00% | New | 4 | $893 |
| TD Private Client Wealth LLC | 0.00% | New | 4 | $893 |
| Clearstead Advisors, LLC | 0.00% | No change | 4 | $820 |
| Sachetta, LLC | 0.00% | No change | 3 | $755 |
| Parkside Financial Bank & Trust | 0.00% | No change | 2 | $448 |
| Manchester Capital Management LLC | 0.00% | No change | 1 | $224 |
| SJS Investment Consulting Inc. | 0.00% | No change | 1 | $224 |
| DiNuzzo Private Wealth, Inc. | 0.00% | No change | 1 | $224 |
| Versant Capital Management, Inc | 0.00% | No change | 1 | $223 |
"% of holder's portfolio" is PEN's weight in each manager's 13F, not the share of Penumbra Inc. the manager owns.